Ray Romano

Manny (voice)

John Leguizamo

Sid (voice)

Denis Leary

Diego (voice)

Queen Latifah

Ellie (voice)

Seann William Scott

Crash (voice)

Josh Peck

Eddie (voice)

Simon Pegg

Buck (voice)

Keke Palmer

Peaches (voice)

Jennifer Lopez

Shira (voice)

Neil deGrasse Tyson

Neil deBuck Weasel (voice)

Jesse Tyler Ferguson

Shangri Llama (voice)

Jessie J

Brooke (voice)

Adam Devine

Julian (voice)

Wanda Sykes

Granny (voice)

Melissa Rauch

Francine (voice)

Michael Strahan

Teddy (voice)

Nick Offerman

Gavin (voice)

Max Greenfield

Roger (voice)

Lilly Singh

Bubbles and Misty (voice)

Chris Wedge

Scrat (voice)

Stephanie Beatriz

Gertie (voice)

Robert Cardone

Freaky Bobby (voice)

Byron Thames

Additional Voices (voice)

Rif Hutton

Additional Voices (voice)

Chris Campbell

Additional Voices (voice)

Holly Dorff

Additional Voices (voice)

Moosie Drier

Additional Voices (voice)

Jason Fricchione

Additional Voices (voice)

Jackie Gonneau

Additional Voices (voice)

Hope Levy

Additional Voices (voice)

Terence Mathews

Additional Voices (voice)

Edie Mirman

Additional Voices (voice)

Juan Pope

Additional Voices (voice)

Ruth Zalduondo

Additional Voices (voice)

David Zyler

Additional Voices (voice)

Ashley Peldon

Additional Voices (voice)

Carlos Ponce

Mariachi Beaver (voice)

Mike Thurmeier

Gravedigger Beaver / Party Molehog (voice)

Randy Thom

Hyrax Minister (voice)

Jorge Lucas

Grandpa Molehog (voice)

James Palumbo

Panicked Start (voice)